Fishes superclass pisces class chondrichthyes subclass elasmobranchii order squaliformes Echinorhinidae — Bramble sharks



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Isla del Coco National Park
Species Lists*

Updated 22 July 2013



Fishes
SUPERCLASS PISCES

CLASS CHONDRICHTHYES


SUBCLASS ELASMOBRANCHII

ORDER Squaliformes

Echinorhinidae — Bramble sharks

Echinorhinus cookei Pietschmann, 1928
ORDER Orectolobiformes

Rhincodontidae — Whale shark

Rhincodon typus Smith, 1828

ORDER Lamniformes

Alopiidae Thresher sharks

Alopias vulpinus (Bonnaterre, 1788)

Odontaspididae - Sand tiger sharks

Odontaspis ferox (Risso, 1810)

ORDER Carcharhiniformes

Carcharhinidae — Requiem sharks

Carcharhinus albimarginatus (Rüppell, 1837)

Carcharhinus falciformis (Müller & Henle, 1839)

Carcharhinus galapagensis (Snodgrass & Heller, 1905)

Carcharhinus limbatus (Müller & Henle, 1839)

Galeocerdo cuvier (Peron & LeSueur in: LeSueur, 1822)

Rhizoprionodon longurio (Jordan & Gilbert, 1882)

Triaenodon obesus (Rüppell, 1837)

Sphyrnidae — Hammerhead sharks

Sphyrna lewini (Griffith & Smith, 1834)

Sphyrna mokarran (Rüppel, 1837)

ORDER Rajiformes

Rhinobatidae — Guitarfishes

Rhinobatos planiceps Garman, 1880

ORDER Myliobatiformes

Dasyatidae — Whiptail stingrays

Taeniura meyeni Müller & Henle, 1841

Myliobatidae — Eagle rays

Aetobatus narinari (Euphrasen, 1790)
Rhinopteridae — Cow-nosed rays

Rhinoptera steindachneri Evermann & Jenkins, 1892

Mobulidae — Mantas, devil rays and mobulas

Manta birostris (Walbaum, 1792)

Mobula munkiana Notarbartolo-di-Sciara, 1987

Mobula tarapacana (Philippi, 1893)

CLASS ACTINOPTERYGII

ORDER Anguilliformes

Myrocongridae — Conger-morays

Myroconger nigrodentatus Castle and Bearez, 1995

Muraenidae — Morays

Anarchias galapagensis (Seale, 1940)

Echidna nebulosa (Ahl, 1789)

Echidna nocturna (Cope, 1872)

Enchelycore octaviana (Myers & Wade, 1941)

Gymnomuraena zebra (Shaw in: Shaw & Nodder, 1797)

Gymnothorax angusticeps (Hildebrand and Barton, 1949)

Gymnothorax buroensis (Bleeker, 1857)

Gymnothorax castaneus (Jordan & Gilbert, 1883)

Gymnothorax dovii (Günther, 1870)

Gymnothorax flavimarginatus (Rüppell, 1830)

Gymnothorax javanicus (Bleeker, 1859)

Gymnothorax meleagris (Shaw in: Shaw & Nodder, 1795)

Gymnothorax panamensis (Steindachner, 1876)

Gymnothorax pictus (Ahl, 1789)

Muraena argus (Steindachner, 1870)

Muraena lentiginosa Jenyns, 1842

Scuticaria tigrina (Lesson, 1828)

Uropterygius macrocephalus (Bleeker, 1865)

Uropterygius versutus Bussing, 1991

Congridae — Conger and garden eels

Ariosoma gilberti (Ogilby, 1898)

Heteroconger klausewitzi (Eibl-Eibesfeldt & Köster, 1983)

Paraconger californiensis Kanazawa, 1961

Ophichthidae — Snake eels

Bascanichthys bascanoides Osburn & Nichols, 1916

Callechelys eristigma McCosker & Rosenblatt, 1972

Gordichthys cf. combibus Lavenberg, ms

Herpetoichthys fossatus (Myers & Wade, 1941)

Ichthyapus selachops (Jordan & Gilbert, 1882)

Myrichthys aspetocheiros McCosker & Rosenblatt, 1993

Myrichthys tigrinus Girard, 1859

Ophichthus rugifer Jordan & Bollman, 1890

Paraletharchus pacificus (Osburn & Nichols, 1916)

Quassiremus evionthas (Jordan & Bollman, 1890)

Quassiremus nothochir (Gilbert, 1890)
ORDER Gonorynchiformes

Chanidae — Milkfishes

Chanos chanos (Forsskål, 1775)

ORDER Aulopiformes

Aulopidae — Flagfins

Aulopus bajacali Parin & Kotlyar, 1984

Chlorophthalmidae — Greeneyes

Chlorophthalmus mento Garman, 1899

Synodontidae — Lizardfishes

Synodus lacertinus Gilbert, 1890

ORDER Ophidiiformes

Ophidiidae — Cusk-eels

Brotula ordwayi Hildebrand & Barton, 1949



Otophidium indefatigabile Jordan & Bollman, 1890

Petrotyx hopkinsi Heller & Snodgrass, 1903

Carapidae — Pearlfishes

Carapus dubius (Putnam, 1874)

Echiodon exsilium Rosenblatt, 1961

Encheliophis vermicularis Müller, 1842

Bythitidae — Viviparous brotulas

Ogilbia cocoensis Möller, Schwarzhans & Nielsen, 2005



ORDER Gadiformes

Moridae - Codlings (Moras)

Physiculus sp.

ORDER Lophiiformes

Lophiidae — Goosefishes

Lophiodes caulinaris (Garman, 1899)

Antennariidae — Frogfishes

Antennarius avalonis Jordan & Starks, 1907

Antennarius coccineus (Lesson, 1831)

Antennarius commerson (Latreille, 1804)

Antennarius sanguineus Gill, 1863

Antennatus strigatus (Gill, 1863)

Ogcocephalidae — Batfishes

Ogcocephalus porrectus Garman, 1899



Zalieutes elater (Jordan & Gilbert, 1882)

ORDER Mugiliformes

Mugilidae — Mullets

Agonostomus monticola (Bancroft, 1834)

Chaenomugil proboscideus (Günther, 1861)

Mugil curema Valenciennes in: Cuvier & Valenciennes, 1836

ORDER Atheriniformes

Atherinopsidae — New World silversides

Atherinella eriarcha Jordan & Gilbert, 1882

Melanorhinus cyanellus (Meek & Hildebrand, 1923)

ORDER Beloniformes

Exocoetidae — Flyingfishes

Exocoetus monocirrhus Richardson, 1846

Exocoetus volitans Linnaeus, 1758

Hirundichthys speculiger (Valenciennes, 1847)

Prognichthys sealei Abe, 1965

Prognichthys tringa Breder, 1928

Hemiramphidae — Halfbeaks

Euleptorhamphus viridis (Van Hasselt, 1823)

Hyporhamphus rosae (Jordan & Gilbert, 1880)

Oxyporhamphus m. micropterus (Valenciennes in: Cuvier & Valenciennes, 1847)
Belonidae — Needlefishes

Platybelone argalus pterura (Osburn & Nichols, 1916)

Strongylura exilis (Girard, 1854)

Tylosurus acus melanotus (Bleeker, 1851)

Tylosurus crocodilus fodiator Jordan & Gilbert, 1882

ORDER Ateleopodiformes

Ateleopodidae — Jelly nose fishes

Guentherus altivela Osório, 1917

ORDER Beryciformes

Holocentridae — Soldierfishes and squirrelfishes

Myripristis berndti Jordan & Evermann, 1903

Myripristis leiognathus Valenciennes, 1846

Plectrypops lima (Valenciennes in: Cuvier & Valenciennes, 1831)

Sargocentron suborbitalis (Gill, 1863)

ORDER Gasterosteiformes

Aulostomidae — Trumpetfishes

Aulostomus chinensis (Linnaeus, 1766)

Fistulariidae — Cornetfishes

Fistularia commersonii Rüppell, 1838

Syngnathidae — Pipefishes and seahorses

Bryx veleronis Herald, 1940

Doryrhamphus excisus excisus Kaup, 1856

Hippocampus ingens Girard, 1858

ORDER Scorpaeniformes

Scorpaenidae — Scorpionfishes

Pontinus clemensi Fitch, 1955

Pontinus strigatus Heller & Snodgrass, 1903

Scorpaena afuerae Hildebrand, 1946

Scorpaena cocosensis Motomura, 2004



Scorpaena histrio Jenyns, 1840

Scorpaena mystes Jordan & Starks in: Jordan, 1895

Scorpaena russula Jordan & Bollman, 1890

DEEP Scorpaenodes rubrivinctus Poss, McCosker & Baldwin, 2010



Scorpaenodes xyris (Jordan & Gilbert, 1882)

Triglidae — Searobins

Bellator farrago Richards & McCosker, 1998
Peristediidae — Armored searobins

Peristedion nesium Bussing, 2010



ORDER Perciformes

Serranidae — Groupers and sea basses

Alphestes immaculatus Breder, 1936

Cephalopholis panamensis (Steindachner, 1877)

Dermatolepis dermatolepis (Boulenger, 1895)

Epinephelus cifuentesi Lavenberg & Grove in: Heemstra & Randall, 1993

Epinephelus itajara (Lichtenstein, 1822)

Epinephelus labriformis (Jenyns, 1840)

Epinephelus mystacinus (Poey, 1852)

Epinephelus niphobles Gilbert & Starks in: Gilbert, 1897

Liopropoma fasciatum Bussing, 1980

Mycteroperca olfax (Jenyns, 1840)

Paranthias colonus (Valenciennes, 1846)

Pronotogrammus eos Gilbert, 1890

Pronotogrammus multifasciatus Gill, 1863

Pseudogramma thaumasium (Gilbert in: Jordan & Evermann, 1900)

Rypticus bicolor Valenciennes, 1846

Serranus aequidens Gilbert, 1890

Serranus tico Meisler & Lavenberg in: Allen & Robertson, 1998



Kuhliidae — Flagtails

Kuhlia mugil (Forster in: Bloch & Schneider, 1801)

Priacanthidae — Bigeyes

Cookeolus japonicus (Cuvier in: Cuvier & Valenciennes, 1829)

Heteropriacanthus cruentatus (Lacépède, 1801)

Pristigenys serrula (Gilbert, 1891)

Apogonidae — Cardinalfishes

Apogon atradorsatus Heller & Snodgrass, 1903

Apogon dovii Günther, 1861

Malacanthidae — Tilefishes

Caulolatilus affinis Gill, 1865

Caulolatilus hubbsi Dooley, 1978

Malacanthus brevirostris Guichenot, 1848

Echeneidae — Suckerfishes, remoras

Echeneis naucrates Linnaeus, 1758

Remora australis (Bennett, 1840)

Remora brachyptera (Lowe, 1839)

Remora osteochir (Cuvier, 1829)

Remora remora (Linnaeus, 1758)

Remorina albescens (Temminck & Schegel, 1850)

Carangidae — Jacks and pompanos

Alectis ciliaris (Bloch, 1787)

Carangoides orthogrammus (Jordan & Gilbert, 1882)

Caranx caballus Günther, 1868

Caranx caninus Günther, 1867

Caranx lugubris Poey, 1860

Caranx melampygus Cuvier in: Cuvier & Valenciennes, 1833

Caranx sexfasciatus Quoy & Gaimard, 1825

Decapterus macarellus (Cuvier in: Cuvier & Valenciennes, 1833)

Elagatis bipinnulata (Quoy & Gaimard, 1825)

Gnathanodon speciosus (Forsskål, 1775)

Naucrates ductor (Linnaeus, 1758)

Selar crumenophthalmus (Bloch, 1793)

Seriola rivoliana Valenciennes in: Cuvier & Valenciennes, 1833

Trachinotus rhodopus Gill, 1863

Trachinotus stilbe (Jordan & McGregor, 1898)

Uraspis helvola (Forster, 1801)

Coryphaenidae — Dolphinfishes

Coryphaena equiselis Linnaeus, 1758

Coryphaena hippurus Linnaeus, 1758

Lutjanidae — Snappers

Aphareus furca (Lacépède, 1801)

Hoplopagrus guntheri Gill, 1862

Lutjanus aratus (Günther, 1864)

Lutjanus argentiventris (Peters, 1869)

Lutjanus guttatus (Steindachner, 1869)

Lutjanus inermis (Peters, 1869)

Lutjanus jordani (Gilbert in: Jordan & Evermann, 1898)

Lutjanus novemfasciatus Gill, 1862

Lutjanus viridis (Valenciennes, 1846)

Gerreidae — Mojarras

Eucinostomus currani Zahuranec in: Yáñez-Arancibia, 1980

Lobotidae — Tripletails

Lobotes pacificus Gilbert, 1898

Haemulidae — Grunts

Anisotremus caesius (Jordan & Gilbert, 1882)

Anisotremus interruptus (Gill, 1862)

Anisotremus taeniatus Gill, 1861

Sciaenidae — Drums

Odontoscion eurymesops (Heller & Snodgrass, 1903)

Umbrina xanti Gill, 1862

Mullidae — Goatfishes

Mulloidichthys dentatus (Gill, 1862)

Kyphosidae — Sea chubs

Kyphosus elegans (Peters, 1869)

Sectator ocyurus (Jordan & Gilbert, 1882)

Chaetodontidae — Butterflyfishes

Chaetodon humeralis Günther, 1860

Chaetodon lunula (Lacépède, 1803)

Johnrandallia nigrirostris (Gill, 1862)

Prognathodes carlhubbsi (Nablant, 1995)

Pomacanthidae — Angelfishes

Holacanthus passer Valenciennes, 1846

Pomacanthus zonipectus (Gill, 1862)

Pomacentridae — Damselfishes

Abudefduf concolor (Gill, 1862)

Abudefduf troschelii (Gill, 1862)

Chromis alta Greenfield & Woods, 1980

Chromis atrilobata Gill, 1862

Microspathodon bairdii (Gill, 1862)

Microspathodon dorsalis (Gill, 1862)

Stegastes acapulcoensis (Fowler, 1944)

Stegastes arcifrons (Heller & Snodgrass, 1903)

Stegastes flavilatus (Gill, 1862)

Stegastes beebei (Nichols, 1924)

Cirrhitidae — Hawkfishes

Cirrhitichthys oxycephalus (Bleeker, 1855)

Cirrhitus rivulatus (Valenciennes, 1846)

Oxycirrhites typus Bleeker, 1857

Sphyraenidae — Barracudas

Sphyraena idiastes Heller & Snodgrass, 1903

Polynemidae — Threadfins

Polydactylus approximans (Lay & Bennett, 1839)

Labridae — Wrasses

Bodianus diplotaenia (Gill, 1862)

Decodon melasma Gomon, 1974

Halichoeres adustus (Gilbert, 1890)

Halichoeres discolor Bussing, 1983



Halichoeres dispilus (Günther, 1864)

Halichoeres nicholsi (Jordan & Gilbert, 1882)

Halichoeres notospilus (Günther, 1864)

Halichoeres salmofasciatus Allen & Robertson, 2002



Iniistius pavo (Valenciennes in: Cuvier & Valenciennes, 1840)

Novaculichthys taeniourus (Lacépède, 1801)

Polylepion cruentum Gomon, 1977

Stethojulis bandanensis (Bleeker, 1851)

Thalassoma grammaticum Gilbert, 1890

Thalassoma lucasanum (Gill, 1862)

Thalassoma purpureum (Forsskål, 1775)

Xyrichtys victori Wellington, 1992

Scaridae — Parrotfishes

Scarus compressus (Osburn & Nichols, 1916)

Scarus ghobban Forsskål, 1775

Scarus perrico Jordan & Gilbert, 1882

Scarus rubroviolaceus Bleeker, 1847

Opistognathidae — Jawfishes

Opistognathus panamaensis Allen & Robertson, 1991

Tripterygiidae — Triplefins

Axoclinus cocoensis Bussing, 1991



Chaenopsidae — Tube blennies

Acanthemblemaria atrata Hastings & Robertson, 1999



Chaenopsis new species

Coralliozetus boehlkei Stephens, 1963

Coralliozetus springeri Stephens & Johnson in: Stephens, Hobson & Johnson, 1966

Emblemaria nivipes Jordan & Gilbert, 1883

Dactyloscopidae — Sand stargazers

Dactyloscopus pectoralis fallax Dawson, 1975

Gillellus chathamensis Dawson, 1977



Labrisomidae — Labrisomid blennies

Dialommus fuscus Gilbert, 1891

Paraclinus mexicanus (Gilbert, 1904)
Blenniidae — Combtooth blennies

Entomacrodus chiostictus (Jordan & Gilbert, 1882)

Hypsoblennius brevipinnis (Günther, 1861)

Ophioblennius steindachneri Jordan & Evermann, 1898

Plagiotremus azaleus (Jordan & Bollman, 1890)

Gobiesocidae — Clingfishes

Arcos rhodospilus (Günther, 1864)

Gobiesox adustus Jordan & Gilbert, 1882

Gobiesox fulvus Meek, 1907

Gobiesox woodsi (Schultz, 1944)

Tomicodon vermiculatus Briggs, 1955



Callionymidae — Dragonets

Synchiropus atrilabiatus (Garman, 1899)

Eleotridae — Sleepers

Eleotris picta Kner, 1863

Eleotris tubularis Heller & Snodgrass, 1903



Gobiidae — Gobies

Bathygobius ramosus longipinnis Ginsburg, 1947

Chriolepis atrimelum Bussing, 1997

Chriolepis dialepta Bussing, 1990

Coryphopterus urospilus Ginsburg, 1938

Elacatinus nesiotes Bussing, 1990

Gobulus hancocki Ginsburg, 1938

Lythrypnus alphigena Bussing, 1990

Lythrypnus cobalus Bussing, 1990

Lythrypnus lavenbergi Bussing, 1990



Lythrypnus rhizophora (Heller & Snodgrass, 1903)

Sicydium cocoensis (Heller & Snodgrass, 1903)



Microdesmidae — Wormfishes

Clarkichthys bilineatus (Clarke, 1936)

Acanthuridae — Surgeonfishes

Acanthurus nigricans (Linnaeus, 1758)

Acanthurus triostegus (Linnaeus, 1758)

Acanthurus xanthopterus Valenciennes in: Cuvier & Valenciennes, 1835

Ctenochaetus marginatus (Valenciennes in: Cuvier & Valenciennes, 1835)

Naso annulatus (Quoy & Gaimard, 1825) vagrant

Naso hexacanthus (Bleeker, 1855) vagrant

Prionurus laticlavius (Valenciennes, 1846)

Zanclidae — Moorish idol

Zanclus cornutus (Linnaeus, 1758)

Gempylidae — Snake mackerels

Gempylus serpens Cuvier, 1829

Scombridae — Mackerels and tunas

Acanthocybium solandri (Cuvier in: Cuvier & Valenciennes, 1832)

Auxis rochei eudorax Collette & Aadland, 1996

Auxis thazard brachydorax Collette & Aadland, 1996

Euthynnus lineatus Kishinouye, 1920

Katsuwonus pelamis (Linnaeus, 1758)

Sarda orientalis (Temminck & Schlegel, 1844)

Scomber japonicus Houttuyn, 1782

Scomberomorus sierra Jordan & Starks in: Jordan, 1895

Thunnus albacares (Bonnaterre, 1788 )

Thunnus obesus (Lowe, 1839)

Istiophoridae — Billfishes

Istiophorus platypterus (Shaw in: Shaw & Nodder, 1792)

Kajikia audax (Philippi, 1887)

Makaira nigricans Lacépède, 1802

ORDER Pleuronectiformes

Bothidae — Lefteye flounders

Bothus leopardinus (Günther, 1862)

Bothus mancus (Broussonet, 1782)

Paralichthyidae — Sand flounders

Hippoglossina bollmani Gilbert, 1890

Syacium latifrons (Jordan & Gilbert, 1882)

Syacium maculiferum (Garman, 1899)



Soleidae — Soles

Aseraggodes herrei Seale, 1940

Cynoglossidae — Tonguefishes

Symphurus varius Garman, 1899

ORDER Tetraodontiformes

Balistidae — Triggerfishes

Balistes polylepis Steindachner, 1876

Canthidermis maculata (Bloch, 1786)

Melichthys niger (Bloch, 1786)

Melichthys vidua (Richardson, 1845)

Pseudobalistes naufragium (Jordan & Starks in: Jordan 1895)

Sufflamen verres (Gilbert & Starks, 1904)

Xanthichthys caeruleolineatus Randall, Matsura & Zama, 1978

Xanthichthys mento (Jordan & Gilbert, 1882)

Monacanthidae — Filefishes

Aluterus monoceros (Linnaeus, 1758)

Aluterus scriptus (Osbeck, 1765)

Cantherhines dumerilii (Hollard, 1854)

Ostraciidae — Boxfishes

Ostracion meleagris Shaw in: Shaw & Nodder, 1796

Tetraodontidae — Puffers

Arothron hispidus (Linnaeus, 1758)

Arothron meleagris (Lacépède, 1798)

Canthigaster punctatissima (Günther, 1870)

Sphoeroides angusticeps (Jenyns, 1842)

Sphoeroides lobatus (Steindachner, 1870)

Diodontidae — Porcupinefishes

Chilomycterus reticulatus (Linnaeus, 1758)

Diodon holocanthus Linnaeus, 1758

Diodon hystrix Linnaeus, 1758
Σ indicates endemic to Isla del Coco

DEEP indicates found very deep


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